Iain, Now that sounds quite intriguing. It also sounds like it might get a bit technical, so perhaps we'll have to continue the discussion off list. The Trillium are interesting, among other reasons, because they seem to interbreed promiscuously wherever their rangers intersect. How real are any of the Trillium species? Are they any older than 10,000-20,000 years, about when the last glaciation ended? The previous interglacial period reached its warmest point at 126,000 years ago. The last glacial age lasted from about 120,000 years ago up until about 12,000 years ago. The peak of ice formation was probably about 20,000 years ago. The ice only reached down to about where I live -- central Indiana. The unglaciated parts of North America may nevertheless have been cold, barren, and relatively dry. Where were all the Trillium species during that ca. 100,000 years of glaciers? I suspect that they were in today's coastal plains of Georgia, Florida, Alabama, and Mississippi; and the coastal plain species were out on the dry continental shelf areas now under up to 200 feet of sea water in the Gulf of Mexico. They may have undergone new speciation processes in the course of surviving the long glacial age far from where they live today. They may be undergoing new speciation at the present, as they encounter each other anew. As I said, we should probably continue off-line. Jim Shields in currently unglaciated central Indiana, but where 20,000 years ago there would have been a mile of ice over my head. At 08:49 PM 5/2/2008 +0100, you wrote: >In his post, # 10, Jim Shields mentions a friend studying possible >relationships amongst Trillium, in particular the, or any, possible >"relationship within or between the yellow Trillium luteum and the >red-brown Trillium cuneatum along with plants of the latter within >populations of the former, posing the question, are they mutants or >hybrids". I would suggest neither. > >If Jim cares to put his friend in touch I think, initially we may have a >three way exchange before I stick my head above the parapet, as I may have >a possible suggestion based on a similar analogue I am researching now >within the genus Lilium. Early days at the moment but I am extremely >intrigued by this situation as it seems to confirm a hypothesis which I am >investigating, no more than an hypothesis quite yet and MIGHT? also have >an implication for the relative relationships between Liliaceae and >Trilliaceae?? > >Iain >_______________________________________________ ************************************************* Jim Shields USDA Zone 5 Shields Gardens, Ltd. P.O. Box 92 WWW: http://www.shieldsgardens.com/ Westfield, Indiana 46074, USA Tel. ++1-317-867-3344 or toll-free 1-866-449-3344 in USA